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新学期开始，各个社团都在大力招新，青春学园的强势社团网球社迎来了一位从美国回来的新生越前龙马（皆川纯子配音）。个头虽小气场却不小的龙马寡言少语性格颇拽，引来了前辈们的不满。但一经较量后，发现龙马果真实力非凡，虽然是一年级，但还是入选为正式队员，目标直指全国大赛。　　而网球社的成员也都各有各的本领和个性。总是笑眯眯的不二（甲斐田雪配音）、爱操心的大石（近藤孝行配音）、龙马的好友桃城（小野坂昌也配音）、桃城的死对头海棠（喜安浩平配音）、闹腾的菊丸（高桥广树配音）、数据控的乾（津田健次郎配音）以及实力强劲的队长手冢（置鲇龙太郎配音）。性格各异的他们在磨合中逐渐形成默契，一路征战，遇见形形色色的来自其他学园的挑战，而大家的目标只有一个，那就是全国大赛的冠军。